Srivastava · FASEB journal : official publication of the Federation of American Societies for Experimental Biology 2012 · controlled animal feeding study · n=?

Mitochondrial biogenesis and increased uncoupling protein 1 in brown adipose tissue of mice fed a ketone ester diet.

Cited 130 times in the scientific literature.

Level 5 - mechanism / opinion, no new human data

Animal research with no human data

PubMed 22362892 · doi:10.1096/fj.11-200410 · record verified 2026-08-30

What was done

Eight-week-old male C57BL/6J mice were fed for up to 1 month with a diet where D-β-hydroxybutyrate-(R)-1,3 butanediol monoester (ketone ester, KE) replaced equicaloric amounts of carbohydrate, with equal fat, protein, and micronutrients. The KE group was fed ad libitum, and control mice were pair-fed to the KE group. Authors evaluated blood D-β-hydroxybutyrate levels, food intake, interscapular brown adipose tissue (IBAT) mitochondrial markers, [(18)F]-fluorodeoxyglucose uptake, plasma leptin, resting and 24-hour energy expenditure, body weight, and insulin sensitivity.

What was found

Blood D-β-hydroxybutyrate in the KE group was 3–5 times higher than reported with high-fat ketogenic diets, and voluntary food intake decreased dose-dependently. IBAT showed doubled electron transport chain proteins, uncoupling protein 1 (UCP1), mitochondrial biogenesis-regulating proteins, and [(18)F]-fluorodeoxyglucose uptake. Plasma leptin increased over 2-fold with elevated sympathetic nervous system activity to IBAT. Resting energy expenditure increased by 14% and the quantitative insulin-sensitivity check index rose by 73%. Total 24-hour energy expenditure and body weight showed no significant difference between groups.

Why it matters

This study shows that oral ketone ester supplementation can stimulate brown fat thermogenic machinery, increase resting metabolic rate, and improve insulin sensitivity markers in a rodent model.

Limits

The study was conducted exclusively in young male mice, precluding direct clinical conclusions in humans. The abstract does not state the sample size (n). Despite the 14% rise in resting metabolic rate, total 24-hour energy expenditure and body weight were unaffected.

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