Bray · The American journal of clinical nutrition 2015 · Randomized controlled trial · n=25

Effect of protein overfeeding on energy expenditure measured in a metabolic chamber.

Cited 64 times in the scientific literature.

Level 2 - randomized trial

Individual randomized controlled trial

PubMed 25733634 · doi:10.3945/ajcn.114.091769 · record verified 2026-08-27

What was done

Twenty-five men and women were randomized to consume approximately 40% excess energy for 56 days from diets containing 5%, 15%, or 25% protein (with excess energy provided as fat in the low-protein group). Twenty-four-hour energy expenditure (24EE) and sleeping energy expenditure (SleepEE) were measured in a metabolic chamber on days 1, 14, and 56 of overfeeding, as well as on day 57 during a return to baseline diet. Skeletal muscle biopsy specimens were collected for metabolic and molecular markers.

What was found

In the low-protein group (excess fat), 24EE and SleepEE did not increase on day 1 of overfeeding, and SleepEE remained unchanged across the study. In groups receiving excess protein, acute increases in 24EE and SleepEE correlated with protein intake (r = 0.50, P = 0.02). Over 8 weeks across all three groups, 24EE correlated with protein intake (r = 0.60, P = 0.004) but not with total energy intake (r = 0.16, P = 0.70). Observed 24EE exceeded predicted values on day 1 (P <= 0.05), day 14 (P = 0.0001), and day 56 (P = 0.0007). Protein and fat oxidation were reciprocally related, and change in fat mass was not related to change in energy expenditure.

Why it matters

This study demonstrates that the acute and sustained rise in metabolic rate during overfeeding is driven specifically by protein intake rather than total caloric excess or fat accumulation.

Limits

The sample size is small (n = 25 split across three arms). The abstract does not provide absolute energy expenditure values in calories, baseline participant demographics, or the specific findings from the skeletal muscle biopsies.

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